State Key Laboratory of Crop Biology, College of Life Science, Shandong Agricultural University, Taian, 271018, Shandong, People's Republic of China. Jinan Academy of Agricultural Sciences, Jinan, 250316, Shandong, People's Republic of China. State Key Laboratory of Crop Biology, College of Life Science, Shandong Agricultural University, Taian, 271018, Shandong, People's Republic of China. chenxs@sdau.edu.cn. State Key Laboratory of Crop Biology, College of Life Science, Shandong Agricultural University, Taian, 271018, Shandong, People's Republic of China. xyunwang@sdau.edu.cn.